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Jonathan Cook February 8th, 2004 02:56 PM

Bull Trout
 
rw wrote in message ...

While there's been a long history of debate about the definition
(the "species problem" in biology), few if any serious biologists would
deny that species are objective, concrete phenomena. This greatly
constrains the possible definitions of "species."

There is a spectrum of opinion.


Debate and opinion, yet the definition is concrete and objective?
That's a non-sequiter for me.

On the far right, so to speak, are people
like Jon who insist on the most rigid and absolute definiton


Yep, I'm a Lumper with a capital L.

This leads to absurdities, like the lions and tigers example.


No more absurd than saying a chihuahua is the same species as a
great dane. As I understand it, it takes a good specialist to be
able to correctly identify a lion skeleton from a tiger skeleton.
Surface coloration and a little different hair growth is hardly
ground for speciation -- else we get back to my human and dog
examples. I'd be glad to declare them subspecies -- like cutts
are to rainbows, but I see no concrete scientific reason to say
they are true species.

The truth is in the middle. Species are "real," but pinning them down with
a simple definition is hard.


Ahh, but we have concrete and objective definitions, don't we?

Fortunately, the definition of "species" isn't left to armchair
taxonomists.

As Darwin wrote to Joseph Hooker: "How painfully true is your remark that
no one has hardly the right to examine the question of species who has not
minutely described many."


Ahh yes, we must uphold the high priesthood of rationalism, the
ivory tower scientists. Us laity just can't grasp the reasoning
it takes to distinguish species. We should just blindy follow
and believe our priests, and if we did the world would be a much
better place. Everything they say is true -- how dare we question
it. If we do, they'll just laugh and call our ideas absurd, or
tell us not to try to be armchair priests.

Jon.

JR February 8th, 2004 03:27 PM

Bull Trout
 
Jonathan Cook wrote (about lions and tigers):

I'd be glad to declare them subspecies -- like cutts
are to rainbows, but I see no concrete scientific reason to say
they are true species.


Your case would be stronger if cutts and rainbows weren't themselves
true species g

The truth is in the middle. Species are "real," but pinning them down with
a simple definition is hard.


Ahh, but we have concrete and objective definitions, don't we?


Yes, but not necessarily simple ones. It's no more than a preconceived
notion (based vaguely perhaps on a common catch-all illustrative
simplification previously used in junior-high science classes) that the
definition of species has to be one that is simply stated and
intuitively obvious. I've had lots of lengthy discussions with folks
over the concept of "species" (or, to refer to another current thread
here, on the pros and cons of hatchery policies in the PNW), only to
have the other person say at some point, "oh, well, you know, I don't
believe in evolution," as though one could have an opinion worth
listening to about either subject without at least a basic understanding
of evolution.

Fortunately, the definition of "species" isn't left to armchair
taxonomists.

As Darwin wrote to Joseph Hooker: "How painfully true is your remark that
no one has hardly the right to examine the question of species who has not
minutely described many."


Ahh yes, we must uphold the high priesthood of rationalism, the
ivory tower scientists. Us laity just can't grasp the reasoning
it takes to distinguish species. We should just blindy follow
and believe our priests, and if we did the world would be a much
better place. Everything they say is true -- how dare we question
it. If we do, they'll just laugh and call our ideas absurd, or
tell us not to try to be armchair priests.


This article by Ernst Mayr is very useful reading (and may be where
Steve found his references to "armchair taxonomists" and Darwin's
correspondence with Hooker):

http://members.aol.com/darwinpage/mayrspecies.htm

JR

Wolfgang February 8th, 2004 04:08 PM

Bull Trout
 

"Jonathan Cook" wrote in message
m...
"Wolfgang" wrote in message

...

[Willi wrote:]

With your definition, some of the choices that need to be made

include:

Actually, those are the exact choices I'm trying to take _out_
of the definition.


I think the fundamental problem with the definition of species is that pesky
little definite article, "the". Various definitions are extremely useful to
more or less worthless depending on circumstance.

Plants......you forgot about plants. How about an apple trunk grafted

onto
pear rootstock and later augmented with plum and cherry branches? (It's
doable.....they're all Rosaceae and graft relatively easily.)


Sorry, you'd have to come up with a better example. Grafting has nothing
to do with reproduction. Can apple tree pollen fertilize pear tree
blossoms? I don't know much about plants, but I doubt it. Doctors have
experimented with pig organs in humans, but that doesn't make us the
same species. (although certain individuals might make us think so :-)


Well, I wasn't so much trying to come up with an example as provide an
obviously absurd end point on a spectrum. Though, actually, with a bit of
thought one should be able to see that it really isn't as absurd as it might
appear at first glance. "Can apple pollen fertilize pear tree blossoms?" I
doubt it too. However I'm not certain. There are a couple of things that
make the question interesting. For one thing, both apples and pears have
long been the objects of careful selection. Importantly, this selection is,
by and large, NOT the sort of selective breeding generally conceeded to
mirror natural selection and which, not so incidentally, played an important
role the evolution of Darwin's theories. The thousands of popular varieties
of apples (and, to a lesser extent, pears) developed over the centuries are
mostly a result of selecting a particular fruit and then grafting the stem
it grew on to a larger supportive structure.....vegetative reproduction, as
opposed to genetic recombination via sex. It is a well known (and mostly
true) maxim that if you plant seeds from your favorite apple you end up with
a tree that produces mostly crab apples, or something very similar. The
same is, of course, true for the other popular fruits in the Rose family.
So, what does the prototypical pear look like? Well, despite thousand of
years of selection, the similarities between apples and pears are still
readily apparent. I'd guess that it looks a lot like the prototypical
apple. Could two distinct (if closely related) species of grass cross-breed
and produce corn? Hm.......nah, probably not.

As far as single-celled organisms at the varying complexity levels go,
I don't know enough to comment.


Varying levels of complexity? Well, a paramecium is certainly anatomically
more complex than a cyanobacterium....um, at the level of the limits of
light microscopy, anyway....but it's a mistake to think that complexity on
that level necessarily corresponds in a linear fashion to the scale of DNA.

Species are a fiction. They can be very useful fictions, but the uses

to
which they are put are not always noble or even justifiable.


I think _that's_ something I can agree with.


Good. I'd much rather someone agree with the point while rejecting an
illustrative example than vice versa. :)

Wolfgang



Willi February 8th, 2004 04:53 PM

Bull Trout
 


Jonathan Cook wrote:

"Wolfgang" wrote in message ...

[Willi wrote:]


With your definition, some of the choices that need to be made include:



Actually, those are the exact choices I'm trying to take _out_
of the definition.



You may be trying but I don't think you'll succeed. I think I understand
where you're coming from because that's how I used to look at it. But,
IMO, the over simplification of using the criteria of "breed and produce
fertile offspring" just opens up a different set of ambiguities. It's
not as clear and concise a definition as it seems on the surface.

If you accept man's intervention to define what breeding and fertile
offspring means (which I assume you do - if you don't I can also make
the other side of the argument) you are left with two choices:

1. To decide how much and what kind of intervention is "allowed"

or

2. Accept that the definition of species will keep changing as man's
technology develops. This, IMO, will lead to some VERY strange
conclusions in the future.

Willi






B J Conner February 8th, 2004 07:42 PM

Bull Trout
 
We had a biologist from ODFG give a dog and pony show on the Fish in the
Deschutes. He said there were two populations of Rainbows. Both the same
species of rainbow. One went to sea and one stayed in the river. The only
way thy could tell them apart was analyzing the layers in the otoliths ( ear
bones ). If the fish had been to sea it had a higer ratio of Strontium to
Calcium. That's looking pretty close.


"Bob Weinberger" wrote in message
...

"Willi" wrote in message
...



That site points to research indicating they can breed and produce
fertile offspring. Sounds like a single species to me. (All other
"scientific" reasons for declaring a new species are, IMO, grounded
only in the scientist's desire for recognition and/or career
advancement ;-)



Question for you West coast guys.

Do Redside Rainbows and Steelhead share any watersheds?

If so do they interbreed?

Willi


The Deschutes is famous for both Redsides and Steelhead. To my knowledge,
they are genetically indistinguishable. Some Steelhead fingerlings never

go
to sea and become resident Redsides and some Redsides go to sea and become
Steelhead. Several other rivers in the vicinity have the same situation.
The reason some rainbows exhibit anadromy, while others in the same system
do not, is a mystery to fish biologists. The Steelhead in the coastal
streams are derived from a different strain of rainbow than the Redsides

of
the interior streams. In most coastal streams virtually all the rainbows
become Steelhead. The resident fish in these streams are cutthroat, though
some of the cuttthroat also exhibit a degree of anadromy - going to the

near
shore salt for periods of 3-6mos. and returning as Searun Cutts of 13-18".
Inch for inch they are better fighters than steelhead.


--
Bob Weinberger
La, Grande, OR

place a dot between bobs and stuff and remove invalid to send email





rw February 9th, 2004 01:57 AM

Bull Trout
 
On 2004-02-08 07:56:42 -0700, (Jonathan Cook) said:

As Darwin wrote to Joseph Hooker: "How painfully true is your remark

that no one has hardly the right to examine the question of species who
has not minutely described many."
Ahh yes, we must uphold the high priesthood of rationalism, the
ivory tower scientists. Us laity just can't grasp the reasoning
it takes to distinguish species. We should just blindy follow
and believe our priests, and if we did the world would be a much
better place. Everything they say is true -- how dare we question
it. If we do, they'll just laugh and call our ideas absurd, or
tell us not to try to be armchair priests.


I assume you wouldn't be able to distinguish between the track of an up
quark from a down quark. Biology is considerably more messy than physics. I
take your point about arguing from authority. I don't like it, either.
Still, the fact is that biologists who spend their entire careers figuring
out incredibly complex taxonomic relationships, and defending their
judgements against the arguments of equally well informed peers, have more
credibility (to me) than do you.

-----------------------------------------------------
Cut "to the chase" for my email address.


Chas Wade February 9th, 2004 03:49 AM

Bull Trout
 
JR wrote:
No, seriously, I mean real evidence. You seem to draw all sorts of
conclusions from talks with "a biologist" or "the guy at the hatchery."


Listen asshole, I'm not a lawyer, and I don't have any legal evidence.

Chas
remove fly fish to reply
http://home.comcast.net/~chas.wade/w...ome.html-.html
San Juan Pictures at:
http://home.comcast.net/~chasepike/wsb/index.html



troutbum_mt February 9th, 2004 06:03 AM

Bull Trout
 
net says...
JR wrote:
No, seriously, I mean real evidence. You seem to draw all sorts of
conclusions from talks with "a biologist" or "the guy at the hatchery."


Listen asshole, I'm not a lawyer, and I don't have any legal evidence.


Prove that you aren't a lawyer! I want to see real evidence. bseg
--
Warren
(use troutbum_mt (at) yahoo to reply via email)
For Conclave Info:
http://www.geocities.com/troutbum_mt...nConclave.html

David Snedeker February 9th, 2004 08:38 AM

Bull Trout
 

"Wolfgang" wrote in message
...
Plants......you forgot about plants. How about an apple trunk grafted

onto
pear rootstock and later augmented with plum and cherry branches? (It's
doable.....they're all Rosaceae and graft relatively easily.)


Interesting. "Relatively easily." "Doable." So you've grafted plum and
cherry branches sucessfully onto an apple trunk, have you? What were the
varieties pray tell?

Dave



Wolfgang February 9th, 2004 12:02 PM

Bull Trout
 

"David Snedeker" wrote in message
...

"Wolfgang" wrote in message
...
Plants......you forgot about plants. How about an apple trunk grafted

onto
pear rootstock and later augmented with plum and cherry branches? (It's
doable.....they're all Rosaceae and graft relatively easily.)


Interesting.


Isn't it?

"Relatively easily."


Yep.

"Doable."


Yep.

So you've grafted plum and
cherry branches sucessfully onto an apple trunk, have you?


Nope.

What were the
varieties pray tell?


My uncle and his cousin both grafted various members of the Rosaceae onto
one another. I have no idea what varieties they used. As they were both of
peasant stock with something less than a high school education, neither
spoke any English on arriving in the U.S., and both spent the better part of
their lives doing manual labor during the daylight hours and drinking
themselves into oblivion in the evenings, I suspect neither of them ever
bothered to learn what varieties they were either. What's more, they
probably didn't know a great deal about current events and I'm certain they
couldn't pass themselves off as authorities on suitable substrates for oil
based paints. Uncle Art DID have a velvet collar on a coat he owned back in
the 50s, though.

Wolfgang




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